An experimental stndy into microstructural effects on short fatigue crack behaviour of 19 stainless steel weld metal smooth specimens during low-cycle fatigue is performed by a so-called ‘effective short fatigue crack criterion’. This material has a mixed microstructure in which it is difficult to distinguish the grains and measure the grain diameter. The columnar grain structure is made up of matrix-rich δ ferrite bands, and the distance between the neighbouring rich δ ferrite bands is an appropriate measure- ment for characterizing this structure. Particularly, the effective short fatigue cracks (ESFCs) always initiate from the bands of δ ferrite in the matrix in the weakest zone on one of the specimen surface zones which is orientated in accordance with the inner or outer surface of welded pipe from which the specimens were machined. These cracks exhibit characteristics of the microstructural short crack (MSC) and the physically small crack (PSC) stages. The average length of the ESFCs at the transition between MSC and PSC behaviour is ≈40μm, while the corresponding fatigue life fraction is ≈0.3 at this transition. Different from previous test observations, the growth rate of the dominant effective short fatigue crack in the MSC stage still shows a decrease with fatigue cycling under the present low-cycle fatigue loading levels. A statistical evolution analysis of the growth rates reveals that the short fatigue crack growth is a damage process that gradually evolves from a non-ordered (chaotic) to a perfectly independent stochastic process, and then to an ordered (history-dependent) stochastic state. Correspondingly, the microstructural effects gradually evolve from a weak effect to a strong one in the MSC stage, which maximizes at the transition point. In the PSC stage, the effects gradually evolve from a strong to weak state. This improves our understanding that the short crack behaviour in the PSC stage is mainly related to the loading levels rather than microstructural effects.